"In reality it's the species that has an evolutionary fitness". Sorry but no. Read "The Selfish Gene" by Richard Dawkins. Selection does NOT act on the level o

"In reality it's the species that has an evolutionary fitness". Sorry but no. Read "The Selfish Gene" by Richard Dawkins. Selection does NOT act on the level of a bee colony, it acts on the level of the individual genes to be found within that colony. So sure some bees are infertile, but they are in service of the genes of the fertile bees. They are in effect enslaved by the genes of the Queen bee for instance. Some types of ants will raid the colonies of other ants, and kidnap their children, bring them back, and the kidnapped children were grow up in service/servitude to the colony which had kidnapped them, and join in on future raids of what was once their home nest. The kidnapped ants are in service to their captors, to the genes of their captors. Say for instance the kidnapped ants were black, and their captors were red(or the other way around), so, even though the kidnapped ants do not have the "red" gene, they are helping to promote the selective advantage of the "red" gene by joining in the hunt for the colony of their captors. It is not a matter of "colony a" being selected for over and above "colony b". It is rather the genes of either colony getting selected for over other genes. All life, from viral to the naked apes like us, are subject to "the tyranny of the selfish gene".