A reasonable article, however they do not use the term "subject" correctly. The subject of their example is not just "the smallest", but all of the following mo

A reasonable article, however they do not use the term "subject" correctly. The subject of their example is not just "the smallest", but all of the following modifiers (adjectives and relative clauses). From "the smallest" to "subunit 6 gene" is all one noun phrase, albeit a complex one. The appropriate advice is therefore to have short subject phrases.