The pool of replicators can prune itself in many ways (indeed, it must), but that doesn't mean that those prunings are having any effect on the pool of genes. T

The pool of replicators can prune itself in many ways (indeed, it must), but that doesn't mean that those prunings are having any effect on the pool of genes. The relative frequency of fertility-relevant alleles is just as likely to remain stable. It's called Hardy Weinberg equilibrium and it's the default state for large populations that are not experiencing predation or expanding into new niches.