With these studies, I always wonder what steps they take to account for paternal uncertainty. On a population scale, I expect virtually all heritable traits to

With these studies, I always wonder what steps they take to account for paternal uncertainty. On a population scale, I expect virtually all heritable traits to correlate more strongly with maternal family history than with paternal family history, simply because some portion of the population are (unbeknownst to them) not actually genetically related to their dads.