I dug through the article and the paper it references, and I can't find any definition of what "Biosphere integrity: Genetic" is supposed to be. The paper refer

I dug through the article and the paper it references, and I can't find any definition of what "Biosphere integrity: Genetic" is supposed to be. The paper references two things under "Biosphere" but neither of them sound like something "Genetic" would refer to. EDIT: Managed to find it, it's on https://www.science.org/doi/10.1126/sciadv.adh2458 https://www.science.org/doi/10.1126/sciadv.adh2458 .