The article argues that the pre-Columbian skeletons in Europe (incl. the ones from the graveyard in Hull) had yaws (a disease causes by another spirochete) rat

The article argues that the pre-Columbian skeletons in Europe (incl. the ones from the graveyard in Hull) had yaws (a disease causes by another spirochete) rather than syphilis. Yaws (at least nowadays) is a tropical disease.