Are there any studies on what, if any, animals respond to line drawings (and to what extent)? In particular, do chimpanzees and other apes closely related to hu

Are there any studies on what, if any, animals respond to line drawings (and to what extent)? In particular, do chimpanzees and other apes closely related to humans? It seems somewhat plausible that “lines-as-edges” was a foot in the door for something that specifically evolved in humans as our ancestors began using paintings and drawings for communication. Maybe it was initially just hijacking edge detection so that some images could be conveyed through drawings, and over time, and that developed into a kind of “grammar” for artificially depicting more nuanced images in media where realism was impossible. This would be similar to how recognizing and interpreting different kinds of basic vocalizations (as many animals can) developed into a much more sophisticated mechanism for developing complex language.