It gets even more complicated when it is hypothesized neurons use spike timings to "compute". > A potentially more powerful way to construct symbols with spike

It gets even more complicated when it is hypothesized neurons use spike timings to "compute". > A potentially more powerful way to construct symbols with spikes is to use spike timing, i.e., to consider exactly when spikes occur within the time window. The maximum number of symbols a neuron can construct with spike timing is limited only by the temporal resolution of the code. In formal terms, the information capacity of spike timing is much higher than the information capacity of spike count. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6665236/ https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6665236/