So they react like most wolves to most of those variables, and therefore the sum total of their behaviour is about the same as other populations of wolves in ar

So they react like most wolves to most of those variables, and therefore the sum total of their behaviour is about the same as other populations of wolves in areas where the variables have similar values? Not sure about terrain and population density in Wisconsin, but at a guess, they probably behave much like wolves in central Sweden or Finland then.