Seems like the relative death rates of tusked vs tuskless would matter. Remember the 32% is a percent potentially on a small number. Say to an extreme 99% of tu

Seems like the relative death rates of tusked vs tuskless would matter. Remember the 32% is a percent potentially on a small number. Say to an extreme 99% of tusked elephants were killed, 99% of tuskless were unharmed. If tuskless is a genetic traight, it takes over very quickly even though total population size is smaller.