Lions seem to laugh, too, and aren't in the list: https://www.youtube.com/watch?v=kyJUQuaECrg https://www.youtube.com/watch?v=kyJUQuaECrg The clip description

Lions seem to laugh, too, and aren't in the list: https://www.youtube.com/watch?v=kyJUQuaECrg https://www.youtube.com/watch?v=kyJUQuaECrg The clip description gets it wrong but some comments recognize it. The laughing bit is near the end. Youtube should be a gold mine for ethologists, maybe not for concrete data but at least for ideas for research.