The paper indicates that the test were done in vivo, in mice models. They also used a synthetic version of a particular compound (melittin), as an adjuvant with

The paper indicates that the test were done in vivo, in mice models. They also used a synthetic version of a particular compound (melittin), as an adjuvant with an existing cancer drug. They also determined how the compound affected those cells. (plasma membranes formed pores, and interferring with cancer-signalling pathways) "The study also found when the venom's main component was combined with existing chemotherapy drugs, it was extremely efficient at reducing tumour growth in mice."