I think the point of the article is that even if you can't make your loop any tighter, there can be benefits to timing your actions so it disrupts your opponent

I think the point of the article is that even if you can't make your loop any tighter, there can be benefits to timing your actions so it disrupts your opponent's loop. That's the fighting example. Fights generally develop a rhythm, where people trade punches. You can get a big advantage by striking when it's not "your turn", even if this means waiting.