The new part of the article is the previously unnoticed role of the epididymis. It turned out that upon entering the epididymis sperm shed a bunch of small RNAs

The new part of the article is the previously unnoticed role of the epididymis. It turned out that upon entering the epididymis sperm shed a bunch of small RNAs(which play a major role in epigenetic expression) and then reaquire small RNA's that reflect more closely the father's environment, later along in the duct.