I'm skeptical that a copyright can be pressed at this point, seeing as no action was taken for years as Pepe memes were created and used on 4chan and elsewhere.

I'm skeptical that a copyright can be pressed at this point, seeing as no action was taken for years as Pepe memes were created and used on 4chan and elsewhere. The creator is on record as having knowledge of their use for quite some time, if I'm remembering correctly.