All the factors you mention are accounted for by researchers that use game theory as the basis for evolutionary modeling (for a primer, check out "Evolutionary

All the factors you mention are accounted for by researchers that use game theory as the basis for evolutionary modeling (for a primer, check out "Evolutionary games on graphs" - https://arxiv.org/abs/cond-mat/0607344 https://arxiv.org/abs/cond-mat/0607344).