I think in this case he's including the suburbs in urban, as contrasted with rural living. In that light urbanization is generally harmful, as rural living tend

I think in this case he's including the suburbs in urban, as contrasted with rural living. In that light urbanization is generally harmful, as rural living tends to have less of an effect on wildlife since individuals are rarely stopping foxes, deer, etc from crossing through their property in the same way that miles of sprawl and deforested lawns do.