http://www.scholarpedia.org/article/Spike-timing_dependent_plasticity http://www.scholarpedia.org/article/Spike-timing_dependent_p... But it kind of requires t

http://www.scholarpedia.org/article/Spike-timing_dependent_plasticity http://www.scholarpedia.org/article/Spike-timing_dependent_p... But it kind of requires the structure of the cortical columns to already be in place - which has happened through genetics/evolution in biology. Biologically plausible backpropagation could also work, but I haven't seen anyone successfully apply it in SNN's