The link between lichen and basidiomycetes was known before this paper, but it was mainly thought that most lichens had none of the basidiomycetes spores, e.g.

The link between lichen and basidiomycetes was known before this paper, but it was mainly thought that most lichens had none of the basidiomycetes spores, e.g. : https://www.anbg.gov.au/lichen/basidiolichens.html https://www.anbg.gov.au/lichen/basidiolichens.html